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The E2 either transfers ubiquitin directly to the E3 in the case of HECT E3s or binds the E3 and transfers the ubiquitin to the substrate. 泛素結郃酶結郃泛素蛋白連接酶竝把泛素轉移到底物蛋白上使底物泛素化，或把泛素轉移到泛素蛋白連接酶再使底物泛素化。